Selective Magnetic Nanoheating: Combining Iron Oxide Nanoparticles for Multi-Hot-Spot Induction and Sequential Regulation

✅ 全文

选择性磁纳米加热:结合氧化铁纳米颗粒实现多热点诱导与顺序调控

作者 Jesús G. Ovejero; Ilaria Armenia; David Serantes; S. Veintemillas‐Verdaguer; Nicoll Zeballos; Fernando López‐Gallego; Cordula Grüttner; Jesús M. de la Fuente; M. P. Morales; Valeria Grazú 期刊 Nano Letters 发表日期 2021 ISSN 1530-6984 DOI 10.1021/acs.nanolett.1c02178 类型 原创研究 (Original Research)

📄 中文摘要 Chinese Abstract

中文
磁性纳米颗粒(MNPs)的远程加热能力,特别是基于氧化铁的纳米颗粒,已在癌症热疗、催化和酶热调控等应用中得到广泛探索。MNPs的一个关键优势是能够在交变磁场(AMF)下产生局部热量,同时保持周围介质的整体温度较低——这对于热敏感的生物过程至关重要。先前的研究表明,MNP表面的局部温度(T_LOC)可通过调节AMF参数(频率和场强)以及MNP特性(尺寸、形状、各向异性)来调控。然而,以往的工作主要集中在单一MNP群体上,缺乏在同一反应器内实现选择性、多热点控制的策略。本研究引入了一种新方法,利用不同氧化铁MNP群体的混合物,实现在单一容器中同时或顺序热激活多种酶促反应。

📋 英文结构化总结 English Structured Summary

全文整理

EN

Background:

The contactless heating capacity of magnetic nanoparticles (MNPs), particularly iron oxide-based ones, has been widely explored in applications such as cancer hyperthermia, catalysis, and enzymatic thermal regulation. A key advantage of MNPs is their ability to generate localized heat under alternating magnetic fields (AMFs) while maintaining a low global temperature in the surrounding medium—critical for thermally sensitive biological processes. Previous studies have demonstrated that local temperature (T_LOC) at the MNP surface can be tuned by adjusting AMF parameters (frequency and field intensity) and MNP properties (size, shape, anisotropy). However, prior work largely focused on single-population MNPs, lacking strategies for selective, multi-hot-spot control within one reactor. This study introduces a novel approach using mixtures of distinct iron oxide MNP populations to enable simultaneous or sequential thermal activation of multiple enzymatic reactions in a single pot.

Methods:

Three monodisperse populations of iron oxide MNPs were synthesized via coprecipitation (CP, ~8.5 nm), thermal decomposition (TD, ~20.2 nm), and oxidative precipitation (OP, ~33.2 nm). These were coated with DMSA, PMAO, or PAA, respectively, to ensure colloidal stability and enable functionalization with Cu²⁺-NTA moieties for site-specific binding of His-tagged fluorescent proteins—superfolder GFP (sGFP) and mCherry RFP—which served as molecular nanothermometers. Local temperature (T_LOC) was inferred from AMF-induced fluorescence loss in these proteins, calibrated against bulk heating experiments. Heating performance was assessed via specific absorption rate (SAR) measurements under varying AMF conditions (96–760 kHz; 5–60 mT). Theoretical modeling of hysteresis losses (HL) was performed using Landau–Lifshitz–Gilbert (LLG) macrospin simulations to predict size- and field-dependent heating behavior.

Results:

Each MNP population exhibited distinct magneto-thermal responses: CP-DMSA showed low but stable SAR across AMF conditions; TD-PMAO achieved high SAR at low-frequency/high-field AMFs; OP-PAA performed best at high-frequency/low-field AMFs. Fluorescence-based nanothermometry confirmed that T_LOC was highly dependent on MNP type and AMF settings. For example, OP-RFP reached 85 ± 5 °C under AMF₁ (100 kHz, 50 mT), while TD-sGFP only reached 25 ± 5 °C under the same conditions. Conversely, under AMF₂ (388 kHz, 10 mT), TD-sGFP heated to ~70 °C, whereas OP-RFP cooled to ~50 °C. Crucially, when TD-sGFP and OP-RFP were mixed in one pot, independent T_LOC readings matched those from individual colloids, demonstrating spatially resolved, non-interfering hot spots.

Data Summary:

SAR values ranged from <75 W/g (CP) to ~430 W/g (OP at 100 kHz, 50 mT). T_LOC measurements showed clear differentiation: CP complexes maintained 50–60 °C regardless of AMF; TD complexes reached up to 70 ± 5 °C under optimal high-frequency/low-field conditions; OP complexes achieved 80–90 °C under low-frequency/high-field AMFs. In mixed colloids, T_LOC differences of up to 60 °C were simultaneously maintained between the two MNP types, with global medium temperature stable at 17 ± 1 °C.

Conclusions:

This work demonstrates, for the first time, the feasibility of generating multiple independently controllable hot spots in a single reactor using tailored iron oxide MNPs and AMF tuning. By combining MNPs with distinct anisotropy and size, and using fluorescent proteins as surface-proximal thermometers, the system enables both simultaneous and sequential thermal regulation of biochemical processes. The approach overcomes limitations of uniform heating in multienzymatic cascades and opens new pathways for precision control in synthetic biology and nanomedicine.

Practical Significance:

This technology enables one-pot multienzymatic reactions where each enzyme operates at its own optimal temperature without thermal interference, improving yield and specificity in biocatalysis. It also holds promise for advanced cancer therapies requiring spatiotemporal control of hyperthermia in heterogeneous tumors, as well as for remotely triggered drug release systems and smart diagnostic platforms leveraging dual-color nanothermometry.

📋 中文结构化总结 Chinese Structured Summary

中文

背景:

磁性纳米颗粒(MNPs)的远程加热能力,特别是基于氧化铁的纳米颗粒,已在癌症热疗、催化和酶热调控等应用中得到广泛探索。MNPs的一个关键优势是能够在交变磁场(AMF)下产生局部热量,同时保持周围介质的整体温度较低——这对于热敏感的生物过程至关重要。先前的研究表明,MNP表面的局部温度(T_LOC)可通过调节AMF参数(频率和场强)以及MNP特性(尺寸、形状、各向异性)来调控。然而,以往的工作主要集中在单一MNP群体上,缺乏在同一反应器内实现选择性、多热点控制的策略。本研究引入了一种新方法,利用不同氧化铁MNP群体的混合物,实现在单一容器中同时或顺序热激活多种酶促反应。

方法:

通过共沉淀法(CP,~8.5 nm)、热分解法(TD,~20.2 nm)和氧化沉淀法(OP,~33.2 nm)合成了三种单分散的氧化铁MNP群体。这些MNP分别用DMSA、PMAO或PAA包覆,以确保胶体稳定性,并实现与Cu²⁺-NTA基团的功能化,用于His标签荧光蛋白——超折叠GFP(sGFP)和mCherry RFP——的特异性结合,这些蛋白作为分子纳米温度计。局部温度(T_LOC)通过AMF诱导的荧光损失推断,并对照整体加热实验进行校准。加热性能通过在不同AMF条件(96–760 kHz;5–60 mT)下的比吸收率(SAR)测量进行评估。使用Landau–Lifshitz–Gilbert(LLG)宏观自旋模拟对磁滞损耗(HL)进行理论建模,以预测尺寸和场依赖的加热行为。

结果:

每种MNP群体表现出不同的磁热响应:CP-DMSA在所有AMF条件下表现出较低但稳定的SAR;TD-PMAO在低频/高场AMF下实现高SAR;OP-PAA在高频/低场AMF下表现最佳。基于荧光的纳米测温法证实,T_LOC高度依赖于MNP类型和AMF设置。例如,OP-RFP在AMF₁(100 kHz,50 mT)下达到85 ± 5 °C,而TD-sGFP在相同条件下仅达到25 ± 5 °C。相反,在AMF₂(388 kHz,10 mT)下,TD-sGFP加热至~70 °C,而OP-RFP冷却至~50 °C。关键的是,当TD-sGFP和OP-RFP在单一容器中混合时,独立的T_LOC读数与单独胶体中的读数相匹配,证明了空间分辨、无干扰的热点。

数据总结:

SAR值范围从<75 W/g(CP)到~430 W/g(OP在100 kHz,50 mT下)。T_LOC测量显示明显差异:CP复合物在所有AMF下保持50–60 °C;TD复合物在最佳高频/低场条件下达到最高70 ± 5 °C;OP复合物在低频/高场AMF下达到80–90 °C。在混合胶体中,两种MNP类型之间的T_LOC差异同时保持高达60 °C,而整体介质温度稳定在17 ± 1 °C。

结论:

本研究首次证明了在单一反应器中利用定制氧化铁MNP和AMF调控产生多个独立可控热点的可行性。通过结合具有不同各向异性和尺寸的MNP,并使用荧光蛋白作为表面邻近温度计,该系统实现了生化过程的同时和顺序热调控。该方法克服了多酶级联反应中均匀加热的局限性,为合成生物学和纳米医学中的精确控制开辟了新途径。

实际意义:

该技术实现了单容器多酶反应,其中每种酶在其最佳温度下运行而不产生热干扰,提高了生物催化中的产率和特异性。它还具有在异质性肿瘤中实现热疗时空控制的先进癌症治疗潜力,以及用于远程触发药物释放系统和利用双色纳米测温的智能诊断平台。

📖 英文全文 English Full Text

EN

pmc Nano Lett Nano Lett 822 acssd nl Nano Letters 1530-6984 1530-6992 pmc-is-collection-domain yes pmc-collection-title ACS AuthorChoice PMC8431726 PMC8431726.1 8431726 8431726 34410726 10.1021/acs.nanolett.1c02178 1 Letter Selective Magnetic Nanoheating: Combining Iron Oxide

Nanoparticles for Multi-Hot-Spot Induction and Sequential Regulation https://orcid.org/0000-0003-3774-6589 Ovejero Jesus G. * † ○ https://orcid.org/0000-0002-2854-2907 Armenia Ilaria ‡ ○ https://orcid.org/0000-0002-3860-2133 Serantes David ∥ https://orcid.org/0000-0002-3015-1470 Veintemillas-Verdaguer Sabino † Zeballos Nicoll ⊥ # https://orcid.org/0000-0003-0031-1880 López-Gallego Fernando ⊥ # Grüttner Cordula ∇ https://orcid.org/0000-0003-1081-8482 de la Fuente Jesús M. ‡ § https://orcid.org/0000-0002-7290-7029 Puerto Morales María del † Grazu Valeria * ‡ § † Institute of Materials Science of Madrid (ICMM-CSIC) , Sor Juana Inés de la Cruz

3, 28049 Madrid, Spain ‡ BioNanoSurf Group, Aragon Nanoscience and Materials Institute (INMA-CSIC-UNIZAR),

Edificio I+D , Mariano Esquillor Gómez, 50018 Zaragoza, Spain § Centro de Investigación Biomédica en Red de Bioingeniería,

Biomateriales y Nanomedicina (CIBER-BBN) , Avenida Monforte de Lemos, 3-5, 28029 Madrid, Spain ∥ Applied

Physics Department and Instituto de Investigacións Tecnolóxicas, Universidade de Santiago de Compostela , 15782 Santiago de Compostela, Spain ⊥ Heterogeneous

Biocatalysis Laboratory, Center for Cooperative Research in Biomaterials (CIC biomaGUNE), Basque Research and Technology

Alliance , Paseo de Miramón 194, 20014 Donostia-San

Sebastián, Spain # IKERBASQUE, Basque Foundation for Science , María Díaz de Haro 3, 48013 Bilbao, Spain ∇ Micromod,

Partikeltechnologie GmbH , Friedrich-Barnewitz-Straße 4, 18119 Rostock, Germany * jesus.g.ovejero@csic.es * vgrazu@unizar.es 19 08 2021 08 09 2021 21 17 389850 7213 7220 03 06 2021 11 08 2021 10 09 2021 13 09 2021 02 04 2024 © 2021 The Authors. Published by American Chemical Society 2021 The Authors https://creativecommons.org/licenses/by/4.0/ Permits the broadest form of re-use including for commercial purposes, provided that author attribution and integrity are maintained ( https://creativecommons.org/licenses/by/4.0/ ). The contactless heating capacity of magnetic nanoparticles (MNPs) has been exploited in fields such as hyperthermia cancer therapy, catalysis, and enzymatic thermal regulation. Herein, we propose an advanced technology to generate multiple local temperatures in a single-pot reactor by exploiting the unique nanoheating features of iron oxide

MNPs exposed to alternating magnetic fields (AMFs). The heating power of the MNPs depends on their magnetic features but also on the intensity and frequency conditions of the AMF. Using a mixture of diluted colloids of MNPs we were able to generate a multi-hot-spot reactor in which each population of MNPs can be selectively activated by adjusting the AMF conditions. The maximum temperature reached at the surface of each MNP was registered using independent fluorescent thermometers that mimic the molecular link between enzymes and MNPs. This technology paves the path for the implementation of a selective regulation of multienzymatic reactions. Hot spot Magnetic nanoparticles Iron oxide Thermal regulation Local temperature Nanothermometry Molecular thermometers Enzymes H2020 Future and Emerging Technologies 10.13039/100010664 829162 Fondo Social de la DGA NA NA European Regional Development Fund 10.13039/501100008530 NA Consejo Superior de Investigaciones Científicas 10.13039/501100003339 PIE-201960E062 Ministerio de Economía y Competitividad 10.13039/501100003329 RED2018-102626-T Ministerio de Economía y Competitividad 10.13039/501100003329 PID2019-109514RJ-100 Ministerio de Economía y Competitividad 10.13039/501100003329 MAT2017-88148-R Ministerio de Economía y Competitividad 10.13039/501100003329 BIO2017-84246-C2-1-R European Commission 10.13039/501100000780 H2020- FETOPEN-RIA 829162 pmc-status-qastatus 0 pmc-status-live yes pmc-status-embargo no pmc-status-released yes pmc-prop-open-access yes pmc-prop-olf no pmc-prop-manuscript no pmc-prop-legally-suppressed no pmc-prop-has-pdf yes pmc-prop-has-supplement yes pmc-prop-pdf-only no pmc-prop-suppress-copyright no pmc-prop-is-real-version no pmc-prop-is-scanned-article no pmc-prop-preprint no pmc-prop-in-epmc yes pmc-license-ref CC BY document-id-old-9 nl1c02178 document-id-new-14 nl1c02178 ccc-price Introduction The growing development of magnetic nanoparticles (MNPs) synthesis methods, especially iron oxide nanoparticles, has boosted their applicability on different fields such as biomedicine, 1 water remediation, 2 , 3 and nanocatalysis, 4 among many other. These materials present unique advantages in terms of contactless manipulation, reusability, and biocompatibility, since iron oxide can be easily digested and integrated by bioorganisms. 5 One of their most interesting features is the possibility of inducing local heat by irradiating them with alternating magnetic fields (AMFs). Besides, the MNPs can be prepared as magnetic colloids thanks to their lack of remanence (superparamagnetic regime) and being easily coated with biological components such as proteins or enzymes. These two features make MNPs ideal biocompatible nanoheaters. The inductive heating power of the MNP colloids has been extensively applied to the thermal treatments of tumor cells 6 and more recently to the regulation of enzymatic and catalytic processes. 7 , 8 In contrast to other catalytic applications, the thermal regulation of an enzymatic activity or protein conformation mediated by MNPs requires an extreme control of the local temperatures achieved on the surface of the nanoheaters. The amount of heat dissipated depends on the MNP composition, size, shape, and aggregation state, but it depends also on the specific conditions (frequency and field) of the AMF applied. 9 Tailoring these parameters, it is possible to optimize the local temperature ( T LOC ) induced in the surface of the MNPs to match different optimal operational temperatures ( T OPT ) of proteins or enzymes attached to their surface. Theoretical and experimental assays have shown that, although the temperature generated at the surface of the MNP can reach up to the boiling point of the media, it decays rapidly at a few nanometers from their surface. 10 − 12 Through the preparation of diluted colloids of MNPs, 13 Armenia et al. demonstrated that, taking advantage of this phenomenon, it is possible to create hot spots in the local environment of the enzymes enhancing their efficiency while maintaining the reactor temperature cold. This seminal work opened the gate to the creation of single-pot multienzymatic reactions operating simultaneously at different optimal temperatures or, alternatively, to the sequential activation of multienzymatic cascades by exploiting the versatility of MNPs as nanoheaters. Currently, such a contactless magnetic heating regulation of enzyme activity has been restricted to the use of a single monodisperse population of MNPs with a homogeneous heating capacity. The idea of combining two MNPs populations with well-differentiated anisotropies to develop a selective system of thermal activation was first described by the theoretical studies of Anikeeva’s group in 2014. 14 They recently applied this principle to the remote activation of heat-sensitive cation channels of kidney cells with outstanding results, 15 and the tremendous potential of this technology can be exploited in many other fields such as tumor therapies. 16 However, none of these studies analyze the specific T LOC induced in the surface of each set of MNPs, which is a critical parameter in the case of biological transformations controlled by the biological activity of proteins including enzymes and an apoptotic induction of tumor cells. 17 To analyze this effect, the use of temperature transducers directly linked to the surface of the MNP provides information about the temperature reached at the active position of the regulated protein during an

AMF activation. The use of fluorescent molecules whose emission intensity depends on the temperature is a frequent strategy for local thermometry 18 , 19 with several technological advantages with respect to other nanothermometry alternatives. 18 , 20 Fluorescent proteins, such as the superfolder Green Fluorescent Protein (sGFP) or m-Cherry Red Fluorescent

Protein (RFP), can be genetically engineered to be tagged with a 6xHis polypetide at their N-terminus in order to resemble a typical site-directed orientation link between enzymes and MNPs functionalized with divalent transition-metal coatings. These proteins suffer an irreversible unfolding denaturation with temperature that leads to a linear loss of fluorescence. 21 , 22 Such linearity makes them interesting thermal probes for nanothermometry in intracellular 23 − 25 and in vivo 26 studies. In this Communication, we demonstrate for the first time that, by using a well-designed toolbox of MNPs with different sizes and shapes, it is possible to generate a multi-hot-spot reactor in which the T LOC may be adjusted by tuning the

AMF conditions. For this aim, we developed a set of iron oxide nanoparticles with core sizes between 8 and 32 nm and different organic and polymeric coatings to create a set of magnetic nanoheaters with different heating powers and different optimum AMF conditions for heat dissipation.

The global heating efficiency of the different cores and coatings was evaluated under AMFs between 5 and 60 mT and frequencies between

96 and 760 kHz. The surface of all these MNPs was engineered with different divalent copper-nitrile acetic acid (Cu 2+ -NTA) moieties, to selectively bind recombinant His-tagged variants of sGFP and RFP through a metal chelate affinity. These fluorescent proteins (FPs) were used as a biomolecular model to determine the maximum local temperature induced at the surface of MNPs when exposed to an AMF.

In this way, we were able to measure and correlate the increment of global and local temperatures induced by the magnetic heating of MNPs and establish a versatile toolbox of magnetic nanoheaters that could match the requirements for a simultaneous or sequential activation of multicompenent biology systems in one pot. Results and Discussion The simplest strategy to modify the heating performance of the

MNPs exposed to AMF is to modify their size. For a certain material, the anisotropy energy of the MNPs grows with the volume of the MNP as E A = K eff V , with K eff and V being the effective anisotropy energy constant and volume of the MNP, respectively. Figure 1 shows the theoretical dependence between the hysteresis losses (HL) in the MNPs and the maximum applied magnetic field ( H MAX ) at 100 and 400 kHz for a system of randomly distributed non-interacting monodisperse magnetite MNPs of increasing sizes at T = 300 K, with their magnetization M⃗ being governed by the stochastic form of the Landau-Lifshitz-Gilbert (LLG) equation (see the Supporting Information for details). 27 Figure 1 LLG macrospin simulations prove that the combination of high frequency–low field (blue square) and low frequency–high field (green square)

AMFs offer an interesting mechanism to select the nanoheating activation.

HLs normalized to the reduced anisotropy ( K u ) for MNPs of different sizes exposed to AMF of increasing H MAX and fixed frequencies of (a) f = 100 and (b) 400 kHz. (b) The curves at 800 kHz were included as dash lines with open symbols. The highlighted sizes and field conditions illustrate how the alternate heat activation could be achieved. It can be observed that, independently of the size, the energy dissipated grows with the H MAX following a sigmoidal dependence. The center and height of the sigmoid scale up with the size of the MNP. These graphs show how the large MNPs requires a higher H MAX to produce a significant heat dissipation, but they achieve a higher dissipation power if the applied field is large enough. It can also be noticed that the saturation value for HL increases with the frequency of AMF, but the inflection points of the sigmoid curves suffer a minimum shifting at high frequencies (800 kHz in dash lines). Comparing the HL of intermediate (20 nm) and large MNPs (30 nm) at low and high frequencies, it is possible to extract a general strategy to choose AMF conditions that invert their heating power (blue and green boxes in Figure 1 ). Furthermore, the reduced anisotropy value ( K u ) presents a certain dependence with the nanoparticle size, which may add further possibilities for fine-tuning the heat release that has not been considered in the present simulations, particularly if working with particle sizes around and below the 10 nm range. 28 , 29 On the one hand, below a certain threshold field ( H MAX = 15 mT for this selection) the MNPs of 30 nm do not transform the magnetic energy into heat losses, whereas the 20 nm

MNPs can reach a theoretical limit of 570 W/g by increasing the AMF frequency to 400 kHz (Supporting Information, Figure S1 ). On the other hand, using an intense (50 mT) and low-frequency AMF (100 kHz) the MNPs of 30 nm result in better nanoheaters than the 20 nm ones, reaching a saturation value for HL equivalent to 430 W/g, while that of the 20 nm MNPs is reduced to 180 W/g. This high field–low frequency versus low field–high frequency (high H-low f vs low H-high f) strategy was first proposed by Anikeeva’s group as an AMF tuning parameter to select which MNPs is activated. 14 Please note that the HL data have been plotted normalized by 2 K u (theoretical maximum for a randomly distributed system) 30 to better illustrate the size effects on the heating performance. The corresponding specific absorption rate (SAR) data are shown in Figure S1 , emphasizing the SAR difference due to the proportionality with frequency. In the light of the theoretical results three monodisperse population of MNPs with average sizes ranging from 8.5 to 33.2 nm were prepared.

To that aim, we performed three different synthesis methods to obtain a homogeneous distribution of MNPs with distribution widths (σ TEM ) below 0.25 ( Table S1 ). Figure 2 shows the transmission electron microscopy (TEM) images of the MNPs obtained by coprecipitation (CP), thermal decomposition (TD), and oxidative precipitation (OP).

The CP synthesis generates spheroidal MNPs with an average size of

8.5 ± 2.0 nm. The MNPs prepared by TD present a larger average diameter ( D TEM = 20.2 ± 4.8 nm) and a multicore structure made of aggregates of smaller nanocrystals.

The largest MNPs were obtained by OP. They present a tetrahedral geometry with an average size of 33.2 ± 7.9 nm. The X-ray diffraction patterns confirm an inverse spinel structure corresponding to magnetite/maghemite in all the cases and the polycrystalline structure of TD-MNPs ( Figure S2 ). To generate a selective activation of a single population of MNP it is important that their average sizes are well-separated and that the widths of the size distributions are small. In this respect, it is of remarkable importance the small overlapping between the size distributions presented in Figure 2 e. Figure 2 Three sets of MNPs were produced with different sizes and geometries. (a) Scheme of MNPs used as thermal regulators and the coatings used for stabilization. TEM pictures of MNPs prepared by (b) CP, (c) TD, and (d) OP. (e) TEM size distribution for the three sets of MNPs.

Red curves indicate the log-normal fitting of the size distribution. (f) Dynamic light scattering intensity curves for the hydrodynamic size of the three sets of MNPs. The three systems were decorated with carboxylic groups in order to improve their colloidal stability and also introduce copper-nitrile acetic acid chelates (Cu 2+ -NTA) moieties onto the MNPs surface to ultimately coordinate the His-tagged fluorescent proteins.

The CP MNPs were coated with a thin layer of dimercaptosuccinic acid (DMSA), whereas the larger MNPs prepared by TD and OP were stabilized with long charged polymers such as poly(maleic anhydride- alt -1-octadecene) (PMAO) and poly(acrylic acid) (PAA), respectively.

These polymers introduce a steric barrier that ensures the colloidal stability of MNPs even when dispersed in saline buffers ( Figure S3 ). Figure 2 f shows the hydrodynamic size of the three systems after a surface coating. The samples present a principal hydrodynamic size of ∼100 nm that suggests the formation of primary aggregates made of a few MNPs during the coating ( Table S1 ). In the case of the TD-PMAO sample, a secondary peak that appears at smaller hydrodynamic sizes indicates the presence of more individually coated MNPs. 31 The proper coating of the

MNPs was confirmed by thermogravimetric analysis, infrared spectroscopy, and Z-potential determination ( Figure S4 ). Interestingly, the high-pressure coating protocol of OP-PAA produced a high-quality thin polymeric coating able to stabilize the MNPs with minimal polymeric content (3.4% of organic mass). The magneto-thermal responses of the three systems were evaluated using quasistatic and AMFs. The hysteresis loops presented in Figure 3 a show the magnetic cycle under quasistatic conditions. All of them present a maximum magnetization at ∼105 ± 2 emu/g Fe , which is consistent with maghemite saturation magnetization, 32 but important differences can be appreciated in the low-intensity field range inset. The CP-DMSA and TD-PMAO samples present a similar coercivity ( H C = 2.5 mT). However, the collective magnetic behavior of the nanocrystals inside the TD-PMAO nanoparticle increases significantly the susceptibility of the cycles. 33 , 34 In the opposite extreme, the hysteresis loop of the OP-PAA sample presents larger coercivity ( H C = 3.75 mT) and smaller susceptibility. Figure 3 Specific magnetic features of each MNP generate a different heating power when exposed to AMFs. (a) Magnetization-field hysteresis loops of MNPs under a quasistatic condition. (inset) The central part of a cycle. (b) SAR of MNPs exposed to AMF of 16 mT at increasing frequencies.

SAR vs H MAX dependence of MNPs exposed to (c) low-frequency AMF (96 kHz) and (d) high-frequency AMF (760 kHz). The different magnetic response between the three samples implies a different heating power when exposed to an AMF. The amount of heat dissipated is generally expressed by an empirical parameter called

SAR that quantifies the amount of heat transmitted to the medium. 9 Figure 3 b shows that, at low-intensity AMF ( H MAX = 16 mT), the TD-PMAO sample generates a larger SAR in the whole range of frequencies studied, whereas the SAR is always the minimum for CP-DMSA. By an analysis of the SAR versus H MAX curves presented in Figure 3 c,d, the sigmoidal dependence predicted by the theoretical models can be identified in the three samples at low and high frequencies. It can be observed that the TD-PMAO and OP-PAA are interesting systems to exploit the selective activation strategy based on a high H–low f versus low f–high H strategy. The T LOC induced by the AMF heating was studied using the above-mentioned MNPs conjugated with two different recombinant his-tagged fluorescent proteins, namely, sGFP and RFP.

These two proteins present a similar β-barrel tertiary structure displayed in Figure 4 a, but their different fluorophore centers generate fluorescence spectra with well-separated emission peaks ( Figure 4 b). 35 , 36 The tertiary structure of both proteins is affected by the temperature suffering the loss of their fluorescence intensity. Besides, Figure 4 c shows that, in both cases, the fluorescence of soluble proteins decays linearly as the temperature increases between

20 and 90 °C. The higher reduction observed in sGFP may be attributed to the higher stability of the resonant chain in RFP. 37 Figure 4 sGFP and m-Cherry RFP were used as a thermal probe of the local temperature in the environment of MNPs. (a) Superimposed representation of a three-dimensional structure of the ternary structure of sGFP and RFP (visualized using Protein Imager 42 ). (b) Intensity reduction of sGFP (green) and RFP (magenta) fluorescence spectra with temperature applied using a global heating source (thermoblock).

Dash lines indicate their respective absorption spectra at 20 °C. (c) Relative fluorescence intensity of free sGFP and RFP proteins at increasing temperatures. Relative fluorescence intensity of sGFP and RFP eluted from the surface of (d) CP, (e) TD, and (f) OP after

5 min of incubation in a thermoblock. Estimated local temperature ( T LOC ) registered from fluorescence loss sGFP and RFP eluted from (g) CP, (h) TD, and (i) OP complexes exposed to different AMF conditions for 5 min. The conjugation of the proteins with inorganic substrates may lead to changes and/or a rigidification of their structure that modifies their temperature stability. 38 To analyze the effect of temperature on the fluorescence intensity of the proteins grafted to the MNPs surface, the fluorescent proteins were eluted in the presence of 0.5 M imidazole and segregated from the MNPs by an ultracentrifugation after the thermal treatments. The fluorescence versus temperature curves presented in Figure 4 d–f for MNP-sGFP and MNP-RFP conjugates reveal a loss of linearity for CP-sGFP and TD-RFP samples, while OP

MNPs fluorescent complexes preserved a linear dependence for the two tested proteins. As expected, the interaction between fluorescent proteins and the MNPs substrates alters its thermal stability in different ways depending on the nature of the coating and the interactions formed at the MNPs-protein interphase during protein binding. Biphasic dependences of the fluorescence with a temperature like those observed for CP-sGFP and TD-RFP are usually observed when the fluorophores present two light-emitting states, 39 as in the case of sGFP and RFP. 36 , 40 The transition between these two states depends on the conformation of the nearest amino acids to the chromophore and may be affected by the interaction with MNP coating. 41 In all the complexes, the immobilization of FPs on the three MNPs drives to less thermally stable protein as observed from the higher slope of the fluorescence versus T curves. Such a reduction in the thermal stability of the proteins may represent an advantage in the case of local nanothermometry. The higher slope of the intensity versus temperature curves translates into a higher sensitivity to the T LOC when used as a thermometer.

The linear and polynomial fittings presented as continuous lines in Figure 4 d–f were used as calibration curves to estimate the maximum T LOC achieved at the protein position during magnetic heating experiments. Table S3 collects the slopes of the linear fittings and the polynomial parameters used for the fitting of nonlinear curves. Figure 4 g–i shows the T LOC registered from the fluorescence of sGFP and RFP after exposing MNPs-sGFP and MNPs-RFP complexes to

AMFs with different conditions of frequency and field for 5 min. The temperature registered in the media remained constant at 17 ±

1 °C through all AMF exposure indicating that the inductive heating was constrained to the local environment of the MNPs due to the low concentrations of the colloids (5 μg Fe /mL). The independent measurements of T LOC obtained from sGFP and RFP nanothermometers conjugated to the three types of MNPs present a significant congruence between them despite the differences observed in their calibration curves. The results probe the robustness and versatility of the thermometric system proposed. Besides, the results obtained from local thermometry are in good agreement with the SAR values presented in Figure 3 , once the specific features of each complex are taken into account. The smallest complexes (CP) generate a T LOC between 50 and 60 °C independently on the AMF conditions applied. The SAR values of these particles are also the smallest (<75 W/g) for the AMF conditions explored, but thanks to the close proximity of the molecular thermometers to the surface of the MNPs they reach a moderate T LOC in every AMF condition. In the case of TD complexes, a little increment of T LOC was registered when exposed to a low-frequency AMF (AMF 1 = 100 kHz to 50 mT). The polymeric coating of this sample introduces a thick spacer between MNPs surface and the molecular thermometer. Only when the AMF conditions are highly favorable (AMF 2 = 388 kHz to 15 mT) does the system reache a T LOC of ∼70 ± 5 °C at the protein position. In the case of OP complexes, the T LOC observed at AMF 1 is between 80 and 90 °C and decays to 50–60 °C for AMF 2 . In this sample, the thin PAA coating implies a closer proximity of the thermometer to the surface of the MNPs. This result highlights the importance of controlling the NTA-His-tag bindings and coatings thicknesses to predict the T LOC induced in the protein position. 13 The potential of MNPs to create a selective heating reactor was evaluated by mixing in a single pot TD-sGFP and OP-RFP complexes.

For this experiment a CP sample was excluded due to its weak dependence of T LOC with the AMF conditions, in the range explored. Figure 5 presents T LOC registered by fluorescence nanothermometry when the mixed suspension is exposed to AMF 1 (100 kHz/50 mT) and AMF 2 (388 kHz/10 mT) conditions. The temperatures registered by each nanothermometer match with those observed in individual colloids ( Figure 4 ), confirming the locality of the heat dissipation processes and thermal independence of each system of MNPs. Figure 5 (a) With a mixture colloid of OP-RFP and

TD-sGFP complexes it is possible to induce multiple hot spots in the same reactor and adjust each T LOC by tuning the AMF conditions. (b) Estimated local temperature ( T LOC ) registered from sGFP and RFP fluorescence in the mixture colloid after

5 min of exposure to AMF 1 = 100 kHz to 50 mT and AMF 2 = 388 kHz to 10 mT. Black dash line indicates the global temperature registered in the medium. (c) SAR registered for individual concentrated colloids (1 mg/mL) of OP-RFP and TD-sGFP at AMF 1 and AMF 2 . Furthermore, the temperatures registered at the two AMF conditions prove that this combination of complexes is suitable to perform a simultaneous multihot-spot and a sequential activation of enzymes.

Using AMF 1 , it is possible to create a T LOC of 25 ± 5 °C at the surface of TD-sGFP and

85 ± 5 °C at the surface of OP-RFP in a reactor that maintains its global temperature at 17 ± 1 °C. In contrast, by with

AMF 2 , the T LOC of TD-sGFP rose to 70 °C, and the T LOC of OP-RFP was reduced to 50 °C. Figure 5 c shows that the T LOC registered in diluted colloids of each kind of MNP correlate with their heating power registered in higher concentrations (1 mg/mL). The use of two independent fluorescent thermal probes for the analysis of multi-hot-spots formed in a pot is a landmark for local nanothermometry.

Dual color fluorescence has been previously used for the analysis of mixtures of biological species to characterize their interactions 43 and is a common protocol in ratiometric fluorescence thermometry. 44 , 45 But, to the best of our knowledge, we pioneer their use to determine the local temperatures induced in a mixture of local nanoheaters activated by a common AMF. In contrast to the common fluorescence microscopy, this approach measures the local temperatures obtaining the fluorescence signal for the whole mixture colloid avoiding any selective imaging bias. 44 , 46 Conclusions Using a clever combination of iron oxide magnetic nanoparticles and local thermal probes based on fluorescent proteins we have proved that it is possible to create both sequential and simultaneous multi-hot-spot conditions with different T LOC in a single pot using different AMF settings. The selection of an adequate combination of magnetic nanoparticles requires a careful control of the magnetothermal properties and homogeneity of magnetic nanoheaters but also a precise control on the arrangement of active proteins on their surface. With diluted colloids, it is possible to heat selectively the environment of the nanoparticles maintaining a low global temperature in the dispersing media. The specific features of the magnetic nanoparticles can be tailored to obtain an optimum heating performance at a specific alternating magnetic field. This technology may create a new paradigm in the regulation of biological molecules such as the creation of one-pot multienzymatic cascades operating at multiple optimal temperatures or being sequentialy activated with different magnetic fields. Supporting Information Available The Supporting Information is available free of charge at https://pubs.acs.org/doi/10.1021/acs.nanolett.1c02178 . Experimental section with theoretical and empirical method details. Complementary data about theoretical calculations of SAR; X-ray diffraction patterns of uncoated MNPs; colloidal parameters and stability of the three MNPs; coating characterization (TGA, FT-IR, and Z-potentials); fluorescent proteins physicochemical paramenters and stability analysis; fitting parameters for thermocalibration curves; hydrodynamic sizes of MNP-FP complexes ( PDF ) Supplementary Material nl1c02178_si_001.pdf Author Contributions ○ (J.G.O. and I.A.) These authors contributed equally. The manuscript was written through contributions of all authors but N.Z. All authors have given approval to the final version of the manuscript. The authors declare no competing financial interest. Acknowledgments This work was funded by the European Commission through the HOTZYMES Project (H2020-FETOPEN-RIA 829162), the Spanish

Ministry of Economy and Competitiveness under Grant No. MAT2017-88148-R (AEI/FEDER, UE) and the PIE-201960E062 project, AEI (BIO2017-84246-C2-1-R project to V.G. and J.M.F., and PID2019-109514RJ-100 to D.S.), Nanotechnology in translational hyperthermia (HIPERNANO) - RED2018-102626-T, and

Fondo Social de la DGA (grupos DGA). Authors acknowledge the use of instrumentation as well as the technical advice provided by the National

Facility ELECMI ICTS, node “Laboratorio de Microscopías

Avanzadas” at the University of Zaragoza. Abbreviations MNPs Magnetic Nanoparticles AMF Alternating Magnetic

Fields TLOC Local temperature HL Hysteresis losses HMAX Maximum

Applied Field HC Coercive field SAR Specific Absorption Rate CP Coprecipitation TD Thermal decomposition OP Oxidative precipitation NTA Nitrile Acetic Acid DMSA Dimercaptosuccinic acid PMAO Poly(maleic anhydride- alt -1-octadecene) PAA Poly(acrylic acid) sGFP superfolder Green Fluorescent Protein RFP m-Cherry Red Fluorescent Protein References Colombo M. ; Carregal-Romero S. ; Casula M. F. ; Gutiérrez L. ; Morales M. P. ; Böhm I. B. ; Heverhagen J. T. ; Prosperi D. ; Parak W. J.

Biological Applications of Magnetic Nanoparticles . Chem. Soc. Rev.

2012 , 41 ( 11 ), 4306 – 4334 . 10.1039/c2cs15337h . 22481569 Gallo-Cordova A. ; Veintemillas-Verdaguer S. ; Tartaj P. ; Mazario E. ; Morales M. d. P. ; Ovejero J. G.

Engineering Iron Oxide Nanocatalysts by a Microwave-Assisted Polyol Method for the Magnetically Induced

Degradation of Organic Pollutants . Nanomaterials 2021 , 11 ( 4 ), 1052 10.3390/nano11041052 . 33924017 PMC8072590 Rivera F. L. ; Recio F. J. ; Palomares F. J. ; Sánchez-Marcos J. ; Menéndez N. ; Mazarío E. ; Herrasti P.

Fenton-like Degradation Enhancement of Methylene Blue Dye with Magnetic Heating Induction . J. Electroanal. Chem.

2020 , 879 , 114773 10.1016/j.jelechem.2020.114773 . Zhang Q. ; Yang X. ; Guan J.

Applications of Magnetic Nanomaterials in Heterogeneous Catalysis . ACS Appl. Nano Mater.

2019 , 2 ( 8 ), 4681 – 4697 . 10.1021/acsanm.9b00976 . Van

De Walle A. ; Kolosnjaj-Tabi J. ; Lalatonne Y. ; Wilhelm C.

Ever-Evolving Identity of Magnetic Nanoparticles within

Human Cells: The Interplay of Endosomal Confinement, Degradation,

Storage, and Neocrystallization . Acc. Chem.

Res.

2020 , 53 ( 10 ), 2212 – 2224 . 10.1021/acs.accounts.0c00355 . 32935974 Rubia-Rodríguez I. ; Santana-Otero A. ; Spassov S. ; Tombácz E. ; Johansson C. ; De La Presa P. ; Teran F. J. ; Morales M. d. P. ; Veintemillas-Verdaguer S. ; Thanh N. T. K. ; Besenhard M. O. ; Wilhelm C. ; Gazeau F. ; Harmer Q. ; Mayes E. ; Manshian B. B. ; Soenen S. J. ; Gu Y. ; Millán Á. ; Efthimiadou E. K. ; Gaudet J. ; Goodwill P. ; Mansfield J. ; Steinhoff U. ; Wells J. ; Wiekhorst F. ; Ortega D.

Whither Magnetic Hyperthermia? A Tentative Roadmap . Materials

2021 , 14 ( 4 ), 706 10.3390/ma14040706 . 33546176 PMC7913249 Marbaix J. ; Mille N. ; Lacroix L. M. ; Asensio J. M. ; Fazzini P. F. ; Soulantica K. ; Carrey J. ; Chaudret B.

Tuning the Composition of FeCo Nanoparticle Heating Agents for Magnetically Induced Catalysis . ACS Appl. Nano Mater.

2020 , 3 ( 4 ), 3767 – 3778 . 10.1021/acsanm.0c00444 . PMC7386363 32743352 Ceylan S. ; Friese C. ; Lammel C. ; Mazac K. ; Kirschning A.

Inductive Heating for Organic Synthesis by Using Functionalized Magnetic Nanoparticles

Inside Microreactors . Angew. Chem., Int. Ed.

2008 , 47 ( 46 ), 8950 – 8953 . 10.1002/anie.200801474 . 18924199 PéRigo E. A. ; Hemery G. ; Sandre O. ; Ortega D. ; Garaio E. ; Plazaola F. ; Teran F. J.

Fundamentals and Advances in Magnetic Hyperthermia . Appl. Phys. Rev.

2015 , 2 ( 4 ), 041302 10.1063/1.4935688 . Guisasola E. ; Baeza A. ; Talelli M. ; Arcos D. ; Moros M. ; De La Fuente J. M. ; Vallet-Regí M.

Magnetic-Responsive Release Controlled by Hot Spot Effect . Langmuir

2015 , 31 ( 46 ), 12777 – 12782 . 10.1021/acs.langmuir.5b03470 . 26536300 Riedinger A. ; Guardia P. ; Curcio A. ; Garcia M. A. ; Cingolani R. ; Manna L. ; Pellegrino T.

Subnanometer Local Temperature Probing and Remotely Controlled Drug Release Based on Azo-Functionalized Iron

Oxide Nanoparticles . Nano Lett.

2013 , 13 ( 6 ), 2399 – 2406 . 10.1021/nl400188q . 23659603 Rodríguez-Rodríguez H. ; Salas G. ; Arias-Gonzalez J. R.

Heat Generation in Single Magnetic Nanoparticles under Near-Infrared Irradiation . J. Phys. Chem. Lett.

2020 , 11 , 2182 – 2187 . 10.1021/acs.jpclett.0c00143 . 32119551 Armenia I. ; GrazúBonavia M.

V. ; De Matteis L. ; Ivanchenko P. ; Martra G. ; Gornati R. ; de la Fuente J. M. ; Bernardini G.

Enzyme Activation by Alternating Magnetic Field: Importance of the Bioconjugation Methodology . J. Colloid

Interface Sci.

2019 , 537 , 615 – 628 . 10.1016/j.jcis.2018.11.058 . 30472637 Christiansen M. G. ; Senko A. W. ; Chen R. ; Romero G. ; Anikeeva P.

Magnetically Multiplexed Heating of Single Domain Nanoparticles . Appl. Phys. Lett.

2014 , 104 ( 21 ), 213103 10.1063/1.4879842 . Moon J. ; Christiansen M. G. ; Rao S. ; Marcus C. ; Bono D. C. ; Rosenfeld D. ; Gregurec D. ; Varnavides G. ; Chiang P. ; Park S. ; Anikeeva P.

Magnetothermal Multiplexing for Selective Remote Control of Cell Signaling . Adv. Funct. Mater.

2020 , 30 ( 36 ), 2000577 10.1002/adfm.202000577 . PMC9075680 35531589 Engelmann U. M. ; Roeth A. A. ; Eberbeck D. ; Buhl E. M. ; Neumann U. P. ; Schmitz-Rode T. ; Slabu I.

Combining Bulk Temperature and Nanoheating Enables Advanced Magnetic Fluid Hyperthermia Efficacy on Pancreatic

Tumor Cells . Sci. Rep.

2018 , 8 ( 1 ), 1 – 12 . 10.1038/s41598-018-31553-9 . 30181576 PMC6123461 Creixell M. ; Bohórquez A. C. ; Torres-Lugo M. ; Rinaldi C.

EGFR-Targeted Magnetic Nanoparticle Heaters Kill Cancer Cells without a Perceptible Temperature

Rise . ACS Nano 2011 , 5 ( 9 ), 7124 – 7129 . 10.1021/nn201822b . 21838221 Qin T. ; Liu B. ; Zhu K. ; Luo Z. ; Huang Y. ; Pan C. ; Wang L.

Organic Fluorescent Thermometers: Highlights from 2013 to 2017 . TrAC, Trends Anal. Chem.

2018 , 102 , 259 – 271 . 10.1016/j.trac.2018.03.003 . Paviolo C. ; Clayton A. H. A. ; Mcarthur S. L. ; Stoddart P. R.

Temperature Measurement in the Microscopic Regime: A Comparison between Fluorescence Lifetime- and Intensity-Based Methods . J. Microsc.

2013 , 250 ( 3 ), 179 – 188 . 10.1111/jmi.12033 . 23521067 Bednarkiewicz A. ; Drabik J. ; Trejgis K. ; Jaque D. ; Ximendes E. ; Marciniak L.

Luminescence Based Temperature Bio-Imaging: Status,

Challenges, and Perspectives . Appl. Phys. Rev.

2021 , 8 ( 1 ), 011317 10.1063/5.0030295 . Moreau M. J. J. ; Morin I. ; Schaeffer P. M.

Quantitative Determination of Protein Stability and Ligand Binding Using a Green Fluorescent Protein Reporter

System . Mol. BioSyst.

2010 , 6 ( 7 ), 1285 – 1292 . 10.1039/c002001j . 20454718 Melnik T. ; Povarnitsyna T. ; Solonenko H. ; Melnik B.

Studies of Irreversible Heat Denaturation of Green Fluorescent Protein by Differential Scanning

Microcalorimetry . Thermochim. Acta 2011 , 512 ( 1–2 ), 71 – 75 . 10.1016/j.tca.2010.09.002 . Okabe K. ; Sakaguchi R. ; Shi B. ; Kiyonaka S. Intracellular Thermometry with

Fluorescent Sensors for Thermal Biology . Pflugers

Archiv European Journal of Physiology ; Springer Verlag , 2018 ; pp 717 – 731 . 10.1007/s00424-018-2113-4 . PMC5942359 29397424 Savchuk O. A. ; Silvestre O. F. ; Adão R. M. R. ; Nieder J. B.

GFP Fluorescence Peak Fraction Analysis Based Nanothermometer for the Assessment of

Exothermal Mitochondria Activity in Live Cells . Sci. Rep.

2019 , 9 ( 1 ), 1 – 11 . 10.1038/s41598-019-44023-7 . 31101860 PMC6525231 Hirsch S. M. ; Sundaramoorthy S. ; Davies T. ; Zhuravlev Y. ; Waters J. C. ; Shirasu-Hiza M. ; Dumont J. ; Canman J. C.

FLIRT:

Fast Local Infrared Thermogenetics for Subcellular Control of Protein

Function . Nat. Methods 2018 , 15 ( 11 ), 921 – 923 . 10.1038/s41592-018-0168-y . 30377360 PMC6295154 Donner J. S. ; Thompson S. A. ; Alonso-Ortega C. ; Morales J. ; Rico L. G. ; Santos S. I. C. O. ; Quidant R.

Imaging of Plasmonic Heating in a Living Organism . ACS Nano

2013 , 7 ( 10 ), 8666 – 8672 . 10.1021/nn403659n . 24047507 García-Palacios J. L. ; Lázaro F. J.

Langevin-Dynamics Study of the Dynamical Properties of Small Magnetic Particles . Phys. Rev. B: Condens.

Matter Mater. Phys.

1998 , 58 , 14937 10.1103/PhysRevB.58.14937 . Demortière A. ; Panissod P. ; Pichon B. P. ; Pourroy G. ; Guillon D. ; Donnio B. ; Bégin-Colin S.

Size-Dependent Properties of Magnetic Iron Oxide Nanocrystals . Nanoscale

2011 , 3 , 225 10.1039/C0NR00521E . 21060937 Tong S. ; Quinto C. A. ; Zhang L. ; Mohindra P. ; Bao G.

Size-Dependent Heating of Magnetic Iron Oxide Nanoparticles . ACS Nano

2017 , 11 ( 7 ), 6808 10.1021/acsnano.7b01762 . 28625045 Conde-Leboran I. ; Baldomir D. ; Martinez-Boubeta C. ; Chubykalo-Fesenko O. ; Del Puerto Morales M. ; Salas G. ; Cabrera D. ; Camarero J. ; Teran F. J. ; Serantes D.

A Single Picture Explains Diversity of Hyperthermia Response of Magnetic Nanoparticles . J. Phys. Chem. C

2015 , 119 ( 27 ), 15698 – 15706 . 10.1021/acs.jpcc.5b02555 . Moros M. ; Pelaz B. ; López-Larrubia P. ; García-Martin M. L. ; Grazú V. ; De La Fuente J. M.

Engineering Biofunctional Magnetic Nanoparticles for Biotechnological Applications . Nanoscale

2010 , 2 ( 9 ), 1746 – 1755 . 10.1039/c0nr00104j . 20676420 Sharifi

Dehsari H. ; Ksenofontov V. ; Möller A. ; Jakob G. ; Asadi K.

Determining Magnetite/Maghemite Composition and Core-Shell Nanostructure from Magnetization Curve for Iron Oxide

Nanoparticles . J. Phys. Chem. C 2018 , 122 ( 49 ), 28292 – 28301 . 10.1021/acs.jpcc.8b06927 . Hugounenq P. ; Levy M. ; Alloyeau D. ; Lartigue L. ; Dubois E. ; Cabuil V. ; Ricolleau C. ; Roux S. ; Wilhelm C. ; Gazeau F. ; Bazzi R.

Iron Oxide Monocrystalline Nanoflowers for Highly Efficient Magnetic Hyperthermia . J. Phys. Chem. C

2012 , 116 ( 29 ), 15702 – 15712 . 10.1021/jp3025478 . Kostopoulou A. ; Brintakis K. ; Vasilakaki M. ; Trohidou K. N. ; Douvalis A. P. ; Lascialfari A. ; Manna L. ; Lappas A.

Assembly-Mediated Interplay of Dipolar Interactions and Surface Spin Disorder in Colloidal Maghemite

Nanoclusters . Nanoscale 2014 , 6 ( 7 ), 3764 – 3776 . 10.1039/C3NR06103E . 24573414 Shu X. ; Shaner N. C. ; Yarbrough C. A. ; Tsien R. Y. ; Remington S. J.

Novel Chromophores and Buried Charges Control Color in MFruits . Biochemistry

2006 , 45 ( 32 ), 9639 – 9647 . 10.1021/bi060773l . 16893165 Wu B. ; Chen Y. ; Müller J. D.

Fluorescence Fluctuation Spectroscopy of MCherry in Living Cells . Biophys. J.

2009 , 96 ( 6 ), 2391 – 2404 . 10.1016/j.bpj.2008.12.3902 . 19289064 PMC2907682 Stepanenko O. V. ; Verkhusha V. V. ; Kazakov V. I. ; Shavlovsky M. M. ; Kuznetsova I. M. ; Uversky V. N. ; Turoverov K. K.

Comparative Studies on the Structure and Stability of Fluorescent Proteins EGFP,

ZFP506, MRFP1, “Dimer2”, and DsRed1† . Biochemistry

2004 , 43 , 14913 10.1021/bi048725t . 15554698 Orrego A. H. ; Romero-Fernández M. ; Millán-Linares M. d. C. ; Pedroche J. ; Guisán J. M. ; Rocha-Martin J.

High Stabilization of Enzymes Immobilized on Rigid Hydrophobic Glyoxyl-Supports: Generation of Hydrophilic Environments on Support Surfaces . Catalysts

2020 , 10 ( 6 ), 676 10.3390/catal10060676 . Guo M. ; Xu Y. ; Gruebele M.

Temperature Dependence of Protein Folding Kinetics in Living Cells . Proc. Natl. Acad. Sci. U. S.

A.

2012 , 109 ( 44 ), 17863 – 17867 . 10.1073/pnas.1201797109 . 22665776 PMC3497798 Brejc K. ; Sixma T. K. ; Kitts P. A. ; Kain S. R. ; Tsien R. Y. ; Ormö M. ; Remington S. J.

Structural Basis for Dual Excitation and Photoisomerization of the Aequorea Victoria Green Fluorescent

Protein . Proc. Natl. Acad. Sci. U. S. A.

1997 , 94 ( 6 ), 2306 – 2311 . 10.1073/pnas.94.6.2306 . 9122190 PMC20083 Stepanenko O. V. ; Stepanenko O. V. ; Kuznetsova I. M. ; Verkhusha V. V. ; Turoverov K. K. Beta-Barrel Scaffold of Fluorescent Proteins. Folding, Stability and Role in Chromophore Formation . In International

Review of Cell and Molecular Biology ; Elsevier Inc. , 2013 ; Vol. 302 , pp 221 – 278 . 10.1016/B978-0-12-407699-0.00004-2 . 23351712 PMC3739439 Tomasello G. ; Armenia I. ; Molla G.

The Protein Imager: A Full-Featured Online Molecular Viewer Interface with Server-Side HQ-Rendering Capabilities . Bioinformatics

2020 , 36 ( 9 ), 2909 – 2911 . 10.1093/bioinformatics/btaa009 . 31930403 Chen Y. ; Tekmen M. ; Hillesheim L. ; Skinner J. ; Wu B. ; Müller J. D.

Dual-Color Photon-Counting Histogram . Biophys. J.

2005 , 88 ( 3 ), 2177 – 2192 . 10.1529/biophysj.104.048413 . 15596506 PMC1305269 Nakano M. ; Arai Y. ; Kotera I. ; Okabe K. ; Kamei Y. ; Nagai T.

Genetically Encoded Ratiometric Fluorescent Thermometer with Wide

Range and Rapid Response . PLoS One 2017 , 12 ( 2 ), e0172344 10.1371/journal.pone.0172344 . 28212432 PMC5315395 Suo H. ; Guo C. ; Li T.

Broad-Scope Thermometry Based on Dual-Color Modulation up-Conversion Phosphor Ba5Gd8Zn4O21:Er3+/Yb3+ . J. Phys. Chem. C

2016 , 120 ( 5 ), 2914 – 2924 . 10.1021/acs.jpcc.5b11786 . Silva P. L. ; Savchuk O. A. ; Gallo J. ; García-Hevia L. ; Bañobre-López M. ; Nieder J. B.

Mapping Intracellular Thermal Response of Cancer Cells to Magnetic Hyperthermia Treatment . Nanoscale

2020 , 12 ( 42 ), 21647 – 21656 . 10.1039/C9NR10370H . 32766635

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中文

# 选择性磁性纳米加热:结合氧化铁纳米颗粒实现多热点诱导与顺序调控

磁性纳米颗粒(MNPs)的非接触加热能力已在癌症热疗、催化和酶热调控等领域得到广泛应用。本文提出了一种先进技术,通过利用氧化铁MNPs在交变磁场(AMFs)下独特的纳米加热特性,在单釜反应器中产生多个局部温度。MNPs的加热功率不仅取决于其磁性特征,还取决于AMF的强度和频率条件。通过使用稀释的MNPs胶体混合物,我们能够构建一个多热点反应器,其中每种MNPs群体可通过调节AMF条件选择性地激活。利用独立的荧光温度计(模拟酶与MNPs之间的分子连接)记录了每种MNP表面达到的最高温度。该技术为多酶反应的选择性调控开辟了道路。

## 引言

磁性纳米颗粒(MNPs)合成方法的快速发展,尤其是氧化铁纳米颗粒,推动了其在生物医学、水修复和纳米催化等众多领域的广泛应用。这些材料在可操控性、可重复使用性和生物相容性方面具有独特优势,因为氧化铁可被生物体轻松消化和整合。其最有趣的特征之一是通过交变磁场(AMFs)辐照诱导局部产热的能力。此外,由于MNPs无剩磁(超顺磁态)且易于被蛋白质或酶等生物组分包覆,可制备为磁性胶体。这两个特征使MNPs成为理想的生物相容性纳米加热器。

MNP胶体的感应加热能力已被广泛应用于肿瘤细胞的热疗,近年来也被用于酶和催化过程的调控。与其他催化应用不同,由MNPs介导的酶活性或蛋白质构象的热调控需要对纳米加热器表面达到的局部温度进行精确控制。热量耗散量取决于MNPs的组成、尺寸、形状和聚集状态,但也取决于所施加AMF的具体条件(频率和场强)。通过调控这些参数,可以优化MNP表面诱导的局部温度(T_LOC),以匹配其表面所附着蛋白质或酶的不同最佳操作温度(T_OPT)。

理论和实验研究表明,尽管MNP表面产生的温度可达到介质沸点,但在距其表面几纳米处会迅速衰减。Armenia等人通过制备稀释的MNP胶体,证明利用这一现象可以在酶的局部环境中产生热点,从而提高酶效率,同时保持反应器整体温度较低。这一开创性工作为创建在单釜中同时在不同最佳温度下运行的多酶反应,或利用MNPs作为纳米加热器的多功能性顺序激活多酶级联反应开辟了道路。

目前,这种非接触磁性加热调控酶活性的方法仅限于使用具有均匀加热能力的单一单分散MNP群体。Anikeeva小组在2014年的理论研究中首次描述了结合两种具有明显不同各向异性的MNP群体来开发选择性热激活系统的想法。他们最近将这一原理应用于肾细胞热敏阳离子通道的远程激活,取得了显著成果,该技术的巨大潜力可在肿瘤治疗等其他许多领域得到利用。

然而,这些研究均未分析每组MNP表面诱导的具体T_LOC,而T_LOC是受蛋白质(包括酶)生物活性控制的生物转化以及肿瘤细胞凋亡诱导的关键参数。为分析这一效应,使用直接连接在MNP表面的温度传感器可提供在AMF激活期间被调控蛋白质活性位点所达到温度的信息。使用发射强度依赖于温度的荧光分子是局部热测量的常用策略,与其他纳米测温替代方案相比具有若干技术优势。

荧光蛋白,如超折叠绿色荧光蛋白(sGFP)或m-Cherry红色荧光蛋白(RFP),可通过基因工程在其N端标记6xHis多肽,以模拟酶与包覆有二价过渡金属的MNPs之间典型的定点取向连接。这些蛋白质随温度发生不可逆的去折叠变性,导致荧光线性损失。这种线性使其成为细胞内和体内研究中纳米测温的有吸引力的热探针。

在本通讯中,我们首次证明,通过使用精心设计的具有不同尺寸和形状的MNPs工具箱,可以构建一个多热点反应器,其中T_LOC可通过调节AMF条件进行调整。为此,我们开发了一组核心尺寸在8至32 nm之间的氧化铁纳米颗粒,具有不同的有机和聚合物包覆层,以创建一组具有不同加热功率和不同最佳散热AMF条件的磁性纳米加热器。在5至60 mT和96至760 kHz的AMF条件下评估了不同核心和包覆层的全局加热效率。

所有这些MNPs的表面均经不同二价铜-次氮基乙酸(Cu²⁺-NTA)基团工程化改造,以通过金属螯合亲和力选择性地结合sGFP和RFP的重组His标签变体。这些荧光蛋白(FPs)被用作生物分子模型,以确定MNPs暴露于AMF时其表面诱导的最大局部温度。通过这种方式,我们能够测量和关联MNPs磁性加热诱导的全局和局部温度增量,并建立一种多功能的磁性纳米加热器工具箱,可满足单釜中多组分生物系统同时或顺序激活的要求。

## 结果与讨论

### 尺寸对加热性能的影响

修改MNPs在AMF下加热性能的最简单策略是改变其尺寸。对于特定材料,MNPs的各向异性能量随MNP体积增长,E_A = K_eff V,其中K_eff和V分别为有效各向异性能量常数和MNP体积。图1显示了MNPs中磁滞损耗(HL)与最大施加磁场(H_MAX)之间的理论依赖性,针对在T = 300 K下随机分布的非相互作用单分散磁铁矿MNPs系统,其磁化强度M⃗由Landau-Lifshitz-Gilbert(LLG)方程的随机形式控制。

图1中的LLG宏自旋模拟证明,高频-低频(蓝方块)和低频-高频(绿方块)AMF的组合提供了一种有趣的纳米加热激活选择机制。图1a和1b分别显示了MNPs在f = 100和400 kHz固定频率下,归一化到约化各向异性(Ku)的HL随H_MAX增加的变化曲线。(b)中以虚线和开符号包含了800 kHz的曲线。突出显示的尺寸和场条件说明了如何实现交替热激活。

可以观察到,无论尺寸如何,耗散能量随H_MAX呈S型依赖增长。S型的中心和高点随MNP尺寸增大而上升。这些图表显示大尺寸MNP需要更高的HMAX才能产生显著的热量耗散,但如果施加的场足够大,它们能达到更高的耗散功率。还可以注意到,HL的饱和值随AMF频率增加而增大,但S型曲线的拐点在高频率下发生最小偏移(800 kHz虚线)。通过比较中等尺寸(20 nm)和大尺寸MNP(30 nm)在低频和高频下的HL,可以提取选择AMF条件以反转其加热功率的一般策略(图1中的蓝框和绿框)。

此外,约化各向异性值(Ku)与纳米颗粒尺寸存在一定依赖性,这为精细调控热量释放提供了进一步的可能性,特别是在10 nm及以下尺寸范围的颗粒中,本模拟未考虑这一点。一方面,在低于某个阈值场(此选择中H_MAX = 15 mT)时,30 nm的MNP不会将磁能转化为热损耗,而20 nm的MNP在将AMF频率增加至400 kHz时可达570 W/g的理论极限。另一方面,使用高强度(50 mT)和低频AMF(100 kHz)时,30 nm的MNP比20 nm的MNP成为更好的纳米加热器,HL饱和值相当于430 W/g,而20 nm MNP的HL饱和值降至180 W/g。

这种高场-低频与低场-高频(高H-低f vs 低H-高f)策略最初由Anikeeva小组提出,作为选择激活哪种MNP的AMF调谐参数。请注意,HL数据已除以2Ku(随机分布系统的理论最大值)作图,以更好地说明尺寸对加热性能的影响。相应的比吸收率(SAR)数据显示在图S1中,强调了由于与频率的比例关系导致的SAR差异。

### 三种MNPs群体的制备与表征

根据理论结果,制备了三种平均尺寸在8.5至33.2 nm范围内的单分散MNP群体。为此,我们采用了三种不同的合成方法,以获得尺寸分布宽度(σ_TEM)低于0.25的均匀MNPs分布(表S1)。图2显示了通过共沉淀法(CP)、热分解法(TD)和氧化沉淀法(OP)获得的MNPs的透射电子显微镜(TEM)图像。

CP合成产生平均尺寸为8.5 ± 2.0 nm的球形MNP。通过TD制备的MNP具有更大的平均直径(D_TEM = 20.2 ± 4.8 nm),并由较小纳米晶体聚集形成多核心结构。最大的MNP通过OP获得,呈现四面体几何形状,平均尺寸为33.2 ± 7.9 nm。X射线衍射图谱确认所有情况下均为对应于磁铁矿/磁赤铁矿的反尖晶石结构,以及TD-MNP的多晶结构(图S2)。

为了实现单一MNP群体的选择性激活,它们的平均尺寸需要充分分离且尺寸分布宽度较小。在这方面,图2e中所示的尺寸分布之间的小重叠具有重要意义。

图2. 制备了三组具有不同尺寸和几何形状的MNPs。(a)用作热调节剂的MNPs及其稳定化包覆层的示意图。通过(b)CP、(c)TD和(d)OP制备的MNPs的TEM图像。(e)三组MNPs的TEM尺寸分布。红色曲线表示尺寸分布的对数正态拟合。(f)三组MNPs流体动力尺寸的动态光散射强度曲线。

三个系统均用羧基修饰以改善其胶体稳定性,并在MNP表面引入铜-次氮基乙酸螯合物(Cu²⁺-NTA)基团,以最终配位His标签荧光蛋白。CP MNP包覆了一层薄薄的二巯基丁二酸(DMSA),而通过TD和OP制备的较大MNP则用长链带电聚合物稳定,分别为聚(马来酸酐-交替-1-十八碳烯)(PMAO)和聚丙烯酸(PAA)。这些聚合物引入空间位阻,确保MNPs即使在分散于盐缓冲液中时也能保持胶体稳定性(图S3)。

图2f显示了表面包覆后三个系统的流体动力尺寸。样品呈现约100 nm的主要流体动力尺寸,表明在包覆过程中形成了由少数MNP组成的一级聚集体(表S1)。在TD-PMAO样品中,在较小流体动力尺寸处出现的次级峰表明存在更多单独包覆的MNP。

通过热重分析、红外光谱和Z电位测定确认了MNP的正确包覆(图S4)。有趣的是,OP-PAA的高压包覆协议产生了高质量的薄聚合物包覆层,能够以最小的聚合物含量(3.4%有机质量)稳定MNP。

### 磁热响应评估

使用准静态和AMF评估了三个系统的磁热响应。图3a所示的磁滞回线显示了准静态条件下的磁循环。所有样品在约105 ± 2 emu/g Fe处呈现最大磁化强度,与磁赤铁矿饱和磁化强度一致,但在低强度场范围(插图)可观察到重要差异。CP-DMSA和TD-PMAO样品呈现相似的矫顽力(H_C = 2.5 mT)。然而,TD-PMAO纳米颗粒内纳米晶体的集体磁性行为显著增加了回线的磁化率。相反,OP-PAA样品的磁滞回线呈现更大的矫顽力(H_C = 3.75 mT)和更小的磁化率。

图3. 每种MNP的特定磁性特征在暴露于AMFs时产生不同的加热功率。(a)准静态条件下MNPs的磁化-场磁滞回线。(插图)回线的中心部分。(b)MNPs在16 mT AMF下随频率增加的SAR。(c)低频AMF(96 kHz)和(d)高频AMF(760 kHz)下MNPs的SAR与H_MAX依赖性。

三个样品之间不同的磁响应意味着在暴露于AMF时具有不同的加热功率。耗散热量的量通常用一个称为SAR的经验参数表示,该参数量化传递到介质的热量。图3b显示,在低强度AMF(H_MAX = 16 mT)下,TD-PMAO样品在整个研究频率范围内产生更大的SAR,而CP-DMSA的SAR始终最小。通过分析图3c和3d中所示的SAR与H_MAX曲线,可以在低频和高频下在三个样品中识别出理论模型预测的S型依赖性。

可以观察到,TD-PMAO和OP-PAA是利用基于高H-低f与低f-高H策略的选择性激活的有趣系统。

### 局部温度的荧光纳米测温研究

使用上述MNPs与两种不同的重组His标签荧光蛋白(即sGFP和RFP)偶联,研究了AMF加热诱导的T_LOC。这两种蛋白质呈现相似的β-桶三级结构(图4a),但其不同的荧光团中心产生具有良好分离发射峰的荧光光谱(图4b)。两种蛋白质的三级结构均受温度影响,导致其荧光强度损失。此外,图4c显示,在这两种情况下,可溶性蛋白质的荧光在20至90°C之间随温度升高而线性衰减。sGFP中观察到的更大降低可归因于RFP中共振链的更高稳定性。

图4. sGFP和m-Cherry RFP被用作MNP环境中局部温度的热探针。(a)sGFP和RFP三级结构的三维结构叠加表示(使用Protein Imager可视化)。(b)使用全局加热源(热块)施加温度时sGFP(绿色)和RFP(品红色)荧光光谱的强度降低。虚线表示它们在20°C下的相应吸收光谱。(c)游离sGFP和RFP蛋白质在升温下的相对荧光强度。从(d)CP、(e)TD和(f)OP表面洗脱的sGFP和RFP在热块中孵育5分钟后的相对荧光强度。从暴露于不同AMF条件5分钟的(g)CP、(h)TD和(i)OP复合物洗脱的sGFP和RFP的荧光损失记录的估计局部温度(T_LOC)。

蛋白质与无机基底的偶联可能导致其结构发生变化或刚性化,从而改变其温度稳定性。为分析温度对接枝到MNP表面的蛋白质荧光强度的影响,在0.5 M咪唑存在下洗脱荧光蛋白,并在热处理后通过超速离心与MNPs分离。图4d-f中所示的MNP-sGFP和MNP-RFP偶联物的荧光与温度曲线揭示了CP-sGFP和TD-RFP样品的线性损失,而OP MNP荧光复合物对两种测试蛋白质保持了线性依赖性。

正如预期的那样,荧光蛋白与MNP基底之间的相互作用以不同方式改变其热稳定性,这取决于包覆层的性质以及蛋白质结合过程中在MNP-蛋白质界面处形成的相互作用。荧光与温度的双相依赖性(如CP-sGFP和TD-RFP中观察到的)通常在荧光团呈现两种发光状态时观察到,如sGFP和RFP的情况。这两种状态之间的转变取决于发色团附近氨基酸的构象,并可能受到与MNP包覆层相互作用的影响。

在所有复合物中,FPs在三种MNPs上的固定导致热稳定性降低,这从荧光与T曲线的更大斜率可以看出。这种蛋白质热稳定性的降低在局部纳米测温方面可能代表一个优势。强度与温度曲线的更大斜率转化为用作温度计时对T_LOC的更高灵敏度。

图4d-f中所示的线性和多项式拟合被用作校准曲线,以估计磁性加热实验期间蛋白质位置达到的最大T_LOC。表S3收集了线性拟合的斜率和用于非线性曲线拟合的多项式参数。

图4g-i显示了将MNP-sGFP和MNP-RFP复合物暴露于不同频率和场条件的AMF 5分钟后,通过sGFP和RFP荧光记录的T_LOC。在整个AMF暴露过程中,介质中记录的温度保持在17 ± 1°C恒定,表明由于胶体浓度低(5 μg Fe/mL),感应加热被限制在MNP的局部环境中。

从与三种类型MNP偶联的sGFP和RFP纳米温度计获得的T_LOC独立测量结果之间存在显著一致性,尽管它们的校准曲线存在差异。结果证明了所提出的测温系统的稳健性和多功能性。此外,局部测温获得的结果与图3中所示的SAR值非常吻合,一旦考虑到每种复合物的具体特征。

最小的复合物(CP)产生50至60°C的T_LOC,与施加的AMF条件无关。这些颗粒的SAR值也是所探索AMF条件中最小的(<75 W/g),但由于分子温度计靠近MNP表面,它们在每个AMF条件下都能达到适中的T_LOC。

在TD复合物的情况下,当暴露于低频AMF(AMF 1 = 100 kHz至50 mT)时,记录到T_LOC略有增加。该样品的聚合物包覆层在MNP表面和分子温度计之间引入了厚间隔物。仅当AMF条件高度有利(AMF 2 = 388 kHz至15 mT)时,系统才能在蛋白质位置达到约70 ± 5°C的T_LOC。

在OP复合物的情况下,在AMF 1下观察到的T_LOC在80至90°C之间,而在AMF 2下降至50-60°C。在该样品中,薄的PAA包覆层意味着温度计更靠近MNP表面。这一结果突出了控制NTA-His标签结合和包覆层厚度以预测蛋白质位置诱导的T_LOC的重要性。

### 多热点选择性加热反应器的评估

通过在一釜中混合TD-sGFP和OP-RFP复合物来评估MNPs创建选择性加热反应器的潜力。由于CP样品的T_LOC对AMF条件的依赖性较弱,因此在本实验中排除了CP样品。

图5显示了当混合悬浮液暴露于AMF 1(100 kHz/50 mT)和AMF 2(388 kHz/10 mT)条件时,通过荧光纳米测温记录的T_LOC。每个纳米温度计记录的温度与在单个胶体中观察到的温度匹配(图4),证实了热量耗散过程的局部性和每个MNP系统的热独立性。

图5.(a)使用OP-RFP和TD-sGFP复合物的混合胶体,可以在同一反应器中诱导多个热点,并通过调节AMF条件调整每个T_LOC。(b)暴露于AMF 1 = 100 kHz至50 mT和AMF 2 = 388 kHz至10 mT 5分钟后,混合胶体中sGFP和RFP荧光记录的估计局部温度(T_LOC)。黑色虚线表示介质中记录的全局温度。(c)在AMF 1和AMF 2下,OP-RFP和TD-sGFP单独浓缩胶体(1 mg/mL)的SAR。

此外,在两种AMF条件下记录的温度证明,这种复合物组合适合执行同时多热点和酶的顺序激活。使用AMF 1,可以在保持全局温度为17 ± 1°C的反应器中,在TD-sGFP表面产生25 ± 5°C的T_LOC,在OP-RFP表面产生85 ± 5°C的T_LOC。相比之下,使用AMF 2,TD-sGFP的T_LOC升至70°C,OP-RFP的T_LOC降至50°C。

图5c显示,每种MNP的稀释胶体中记录的T_LOC与其在较高浓度(1 mg/mL)下记录的加热功率相关。使用两种独立的荧光热探针分析釜中形成的多热点是局部纳米测温的一个里程碑。双色荧光先前已被用于分析生物物种混合物以表征其相互作用,并且是比率荧光测温的常见方案。但据我们所知,我们率先使用它们来确定由共同AMF激活的局部纳米加热器混合物中诱导的局部温度。与常见的荧光显微镜不同,这种方法通过获取整个混合胶体的荧光信号来测量局部温度,避免了任何选择性成像偏差。

## 结论

通过使用氧化铁磁性纳米颗粒和基于荧光蛋白的局部热探针的巧妙组合,我们证明了在单釜中使用不同的AMF设置可以创建具有不同T_LOC的顺序和同时多热点条件。选择合适的磁性纳米组合需要仔细控制磁性纳米加热器的磁热性能和均一性,还需要精确控制活性蛋白质在其表面的排列。

使用稀释的胶体,可以选择性地加热纳米颗粒的环境,同时保持分散介质中的低全局温度。可以定制磁性纳米颗粒的特征,以在特定交变磁场下获得最佳加热性能。这项技术可能在生物分子调控中创造一个新的范式,例如创建在多种最佳温度下运行的单釜多酶级联反应,或通过不同的磁场顺序激活。

## 支持信息

支持信息可在https://pubs.acs.org/doi/10.1021/acs.nanolett.1c02178免费获取。包含理论和实验方法细节的实验部分;SAR理论计算的补充数据;未包覆MNPs的X射线衍射图谱;三种MNPs的胶体参数和稳定性;包覆层表征(TGA、FT-IR和Z电位);荧光蛋白理化参数和稳定性分析;热校准曲线的拟合参数;MNP-FP复合物的流体动力尺寸。

## 作者贡献

○(J.G.O.和I.A.)这两位作者贡献均等。手稿由所有作者共同撰写,但N.Z.除外。所有作者均已批准手稿的最终版本。作者声明无竞争性经济利益。

## 致谢

本研究由欧盟委员会通过HOTZYMES项目(H2020-FETOPEN-RIA 829162)、西班牙经济与竞争力部通过MAT2017-88148-R资助号(AEI/FEDER, UE)和PIE-201960E062项目、AEI(V.G.和J.M.F.的BIO2017-84246-C2-1-R项目,D.S.的PID2019-109514RJ-100项目)、转化热疗中的纳米技术(HIPERNANO)-RED2018-102626-T以及DGA社会基金(DGA小组)资助。作者感谢国家设施ELECMI ICTS提供的仪器使用和技术建议,节点为萨拉戈萨大学"先进显微镜实验室"。

## 缩写

MNPs:磁性纳米颗粒;AMF:交变磁场;T_LOC:局部温度;HL:磁滞损耗;H_MAX:最大施加场;H_C:矫顽场;SAR:比吸收率;CP:共沉淀;TD:热分解;OP:氧化沉淀;NTA:次氮基乙酸;DMSA:二巯基丁二酸;PMAO:聚(马来酸酐-交替-1-十八碳烯);PAA:聚丙烯酸;sGFP:超折叠绿色荧光蛋白;RFP:m-Cherry红色荧光蛋白。